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To replace read symbol or gap in the Sanger Reads Editor select the nucleotide and press Shift+R or Edit–>Replace character/gap context menu for switching the edit mode on. 

 While the selection is blinking, press a keyboard character that you want to replace. 

 After that the edit mode is switched off. For replace another symbol press Shift+R again.

 You always can cancel the changes pressing Ctrl+Z.

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